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Godparents to Peter and Alison's daughter Sophie, the mismatched pair are unexpectedly thrown back together under tragic circumstances, when their friends are killed in an accident. The couple had specified that Holly and Messer should be Sophie's guardians in the event of their deaths, so obliged to move in together, ambitious bakery owner Holly and laddish TV sports director Messer find their new arrangement taking a toll on their careers and love lives. The challenges of caring for Sophie manage to bring her surrogate parents closer together, but will hunky paediatrician Sam (Josh Lucas) get in the way of a happy family?